This is only for a veterinarian or dog psychologist: we know dogs have incredibly sensitive noses, which allow them to decode many clues about the outside world. Is it possible that Lola upon being in a forest for the first time smelled the scents of big animals (deer, bear, mountain lion) with which she was not familiar, and therefore she became kind of generally scared?
A lifelong friend of mine who brings his dog to stay with him for weeks at a time in the forest reported: "When I take my dog out to my woods and we haven't been there for a while, he just wants to stay in the car for a first few hours. I think they can sense predators who've been there. At my place that means bear, coyotes and cats."
So it seems plausible that Lola was just nervous from sensing predators. (There are piles of bear poo and deer poo on the property.)